Six months have passed since the victorious Battle of Britannia and therefore the reclamation of Gallia from Neuroi invaders. Yoshika Miyafuji, member of the famed 501st Joint Fighter Wing, has come home to Fuso and graduated from secondary school .
However, her fight is way from over. She receives a letter supposedly sent by her long-deceased father, containing blueprints of a state-of-the-art Striker Unit he had been performing on before his death. The Unit, designed specifically for Yoshika, could be capable of harnessing her extraordinary magical powers.
Meanwhile, a replacement threat in Europe is rising. A Neuroi nest of an unprecedented size and might has appeared over Venezia, wiping out local Witch forces and instantly swallowing the northern a part of the country. to form matters worse, a newly spotted humanoid sort of Neuroi is attempting to return into contact with humans.